Journal Articles and Refereed Papers:



  • Farcuh, M., Copes, B., Le-Navenec, G., Marroquin, J., Jaunet, T., Chi-Ham, C., Cantu, D., Bradford, K.J., Van Deynze, A. (2020) Texture diversity in melon (Cucumis melo L.): Sensory and physical assessments. Postharvest Biology and Technology 159: 111024;
  • Liu, S., Bradford, K.J., Huang, Z., Venable, L. 2020. Hydrothermal sensitivities of seed populations underlie fluctuations of dormancy states in an annual plant community. Ecology 100(3): e02958.


  • Liu, S., Streich, J., Borevitz, J.O., Rice, K.J., Li, T., Li, B., Bradford, K.J. 2019. Environmental resource deficit may drive the evolution of intraspecific trait variation in invasive plant populations. Oikos 128: 171-184. doi:10.1111/oik.05548.


  • Bradford, K.J. 2018. Interpreting biological variation: seeds, populations and sensitivity thresholds. Seed Sci Res 28: 158-167.
  • Bradford, K.J., Dahal, P., Van Asbrouck, J., Kunusoth, K., Bello, P., Thompson, J., Wu, F. 2018. The dry chain: reducing postharvest losses and improving food safety in humid climates. Trends in Food Science & Technology 71: 84-93.
  • Bertier, L., Ron, M., Huo, H., Bradford, K.J., Michelmore, R.W. 2018. High-resolution analysis of the efficiency, heritability, and editing outcomes of CRISPR/Cas9-induced modifications of NCED4 in lettuce (Lactuca sativa). G3: Genes  Genomes Genetics 8: 1513-1521.

  • Timsina, K.P., Bradford, K.J., Dahal, P., Shivakoti, G.P., Kunusoth, K., Van Asbrouck, J., Bajracharya, J., Pandey, I.R. 2018. Potential impacts of desiccant-based drying and hermetic storage on the value chain for onion seeds in Nepal. J. Agribusiness in Developing and Emerging Economies. 8(2): 363-390.


  • Vegh, A., Incze, N., Fábián, A., Huo, H., Bradford, K.J., Balázs, E., Soós, V. 2017. Comprehensive analysis of DWARF14- LIKE2 (DLK2) reveals its functional divergence from strigolactone-related paralogs. Frontiers in Plant Science 8: 1641.


  • Bradford, K.J., Dahal, P., Bello, P. (2016) Using relative humidity indicator paper to measure seed and commodity moisture contents. Agricultural and Environmental Letters 1: 160018. doi:10.2134/ael2016.04.0018.
  • Huo, H., Henry, I.M., Coppoolse, E.R., Verhoef-Post, M., Schut, J.W., de Rooij, H., Vogelaar, A., Joosen, R.V.L., Woudenberg, L., Comai, L., Bradford, K.J. 2016. Rapid identification of lettuce seed germination mutants by bulked segregant analysis and whole genome sequencing. Plant Journal 88: 345-360. DOI: 10.1111/tpj.13267.
  • Bello, P., Bradford, K.J. 2016. Single-seed oxygen consumption measurements and population-based threshold models link respiration and germination rates under diverse conditions. Seed Science Research 26: 199-221.
  • Huo, H., Wei, S., Bradford, K.J. 2016. DELAY OF GERMINATION1 (DOG1) regulates both seed dormancy and flowering time through microRNA pathways. Proc. Natl. Acad. Sci. USA 115: E2199-E2206.
  • Yoong, F.-Y., O’Brien, L.K., Truco, M.J., Huo, H., Sideman, R., Hayes, R., Michelmore, R.W., Bradford, K.J. 2016. Genetic variation for thermotolerance in lettuce seed germination is associated with temperature-sensitive regulation of ETHYLENE RESPONSE FACTOR1 (ERF1). Plant Physiology 170: 472-488.
  • Huang, Z., Liu, S., Bradford, K.J., Huxman, T.E., Venable, D.L. 2016. The contribution of germination functional traits to population dynamics of a desert plant community. Ecology 97: 250-261.


  • Donohue, K., Burghardt, L.T., Runcie, D., Bradford, K.J., and Schmitt, J. 2015. Applying developmental threshold models to evolutionary ecology. Trends in Ecology and Evolution 30: 66–77.
  • Timsina, K.P., Shivakoti, G.P., Bradford, K.J. 2015. Supply situation of vegetable seeds in Nepal: An analysis from policy perspective. Nepalese Horticulture 10: 26-36.


  • Bradford, K.J., Bello, P., Fu, J.C., and Barros, M. 2013. Single-seed respiration: a new method to assess seed quality. Seed Sci. Technol. 41: 420-438.
  • Boddy, L.G., Bradford, K.J., and Fischer, A.J. 2013. Stratification requirements for seed dormancy alleviation in a wetland weed. PLOS One 8(9) e71457. doi:10.1371/journal.pone.0071457
  • Huo, H., Dahal, P., Kunusoth, K., McCallum, C.M., Bradford, K.J. 2013. Expression of 9-cis-EPOXYCAROTENOID DIOXYGENASE 4 (NCED4) is essential for thermoinhibition of lettuce seed germination but not for seed development or stress tolerance. Plant Cell 25: 884-900.


  • Boddy, L.G., Bradford, K.J., and Fischer, A.J. 2012. Population-based threshold models describe weed germination and emergence patterns across varying temperature, moisture and oxygen conditions. J. Applied Ecol. 49: 1225-1236.


  • Schwember, A.R., and Bradford, K.J. 2011. Oxygen interacts with priming, moisture content and temperature to affect the longevity of lettuce and onion seeds. Seed Sci. Res. 21: 175-185.
  • Argyris, J., Truco, M.J., Ochoa, O., McHale, L., Dahal, P., Van Deynze, A., Michelmore, R.W., Bradford, K.J. 2011. A gene encoding an abscisic acid biosynthetic enzyme (LsNCED4) collocates with the high temperature germination locus  Htg6.1 in lettuce (Lactuca sp.). Theor. Appl. Genet. 122: 95-108.
  • Van Deynze, A.E., Hutmacher, R.B., and Bradford, K.J. 2011. Gene flow between Gossypium hirsutum and G. barbadense is asymmetric. Crop Sci. 51: 298-305.


  • Bradford, K.J., and Harada, J.J. 2010. Introduction to Translational Seed Biology: From Model Systems to Crop Improvement. Plant Sci. 179: 553-553. [Introduction to special issue of this journal edited by KJB and J.J. Harada based on Plant Science Symposium we organized in 2007.]
  • Schwember, A.R. and Bradford, K.J. 2010. Quantitative trait loci associated with longevity of lettuce seeds under  conventional and controlled deterioration storage conditions. J. Exp. Bot.: 61: 4423-4436.
  • Miller, J.K., Herman, E.M., Jahn, M., and Bradford, K.J. 2010. Strategic research, education and policy goals for seed science and crop improvement. Plant Sci.: 179: 645–652.
  • Miller, J.K., Bradford, K.J. 2010. The regulatory bottleneck for biotech specialty crops. Nature Biotechnol. 10: 1012-1014.
  • Schwember, A.R., and Bradford, K.J. 2010. A genetic locus and gene expression patterns associated with the priming effect on lettuce seed germination at elevated temperatures. Plant Mol. Biol. 73: 105-118.


  • Argyris, J.M., Dahal, P., Hayashi, E., Still, D.W., and Bradford, K.J. 2008. Genetic variation for lettuce seed thermoinhibition is associated with temperature-sensitive expression of abscisic acid, gibberellin and ethylene biosynthesis, metabolism and response genes. Plant Physiol. 148: 926-947.
  • Bradford, K.J. 2008. Shang Fa Yang: Pioneer in plant ethylene biochemistry. Plant Science 175: 2-7.
  • Bradford, K.J., Benech-Arnold, R., Côme, D., and Corbineau, F. 2008. Quantifying the sensitivity of barley seed germination  to oxygen, abscisic acid and gibberellin using a population-based threshold model. J. Exp. Bot. 59: 335-347.
  • Moravec, C.M., Bradford, K.J., and Laca, E.A. 2008. Water relations of drumstick tree seed (Moringa oleifera): imbibition, desiccation, and sorption isotherms. Seed Sci. & Technol. 36: 311-324.
  • Hill, H., Bradford, K.J., Cunningham, J., and Taylor, A.G. 2008. Primed lettuce seeds exhibit increased sensitivity to moisture during aging. Acta Horticulturae 782: 135-141.


  • Argyris, J., Dahal, P., Truco, M.J., Ochoa, O., Still, D.W., Michelmore, R.W., and Bradford, K.J. 2007. Genetic analysis of lettuce seed thermoinhibition. Acta Hortic. 782: 23-33.
  • Gealy, D.R., Bradford, K.J., Hall, L., Hellmich, R., Raybould, A., Wolt, J., and Zilberman, D. 2007. Implications of Gene Flow  in the Scale-up and Commercial Use of Biotechnology-derived Crops: Economic and Policy Considerations. Council for Agricultural Science and Technology (CAST), Issue Paper 37. CAST, Ames, Iowa.
  • Hill, H.J., Cunningham, J.D., Bradford, K.J., and Taylor, A.G. 2007. Primed lettuce seeds exhibit increased sensitivity to moisture content during controlled deterioration. HortScience 42: 1436-1439.
  • Kalaitzandonakes, N., Alston, J.M., and Bradford, K.J. 2007. Compliance costs for regulatory approval of new biotech crops. Nature Biotech. 25: 509-511.
  • Bradford, K.J., Côme, D., and Corbineau, F. 2007. Quantifying the oxygen sensitivity of seed germination using a population- based threshold model. Seed Sci. Res. 17: 33-43.


  • Alston, J.M., Bradford, K.J., and Kalaitzandonakes, N. 2006. The economics of horticultural biotechnology. J. Crop Improvement 18: 413-431.


  • Argyris, J., Truco, M.J., Ochoa, O., Knapp, S.J., Still, D.W., Lenssen, G.M., Schut, J.W., Michelmore, R.W., and Bradford, K.J. 2005. Quantitative trait loci associated with seed and seedling traits in Lactuca. Theor. Appl. Genet. 111: 1365-1376.
  • Gandhi, S., Heesacker, A., Freeman, C., Argyris, J., Bradford, K.J., Knapp, S.J. 2005. The self-incompatibility locus (S) and quantitative trait loci for self-pollination and seed dormancy in sunflower. Theor. Appl. Genet. 111: 619-629.
  • Van Deynze, A.E., Sundstrom, F.J., and Bradford, K.J. 2005. Pollen-mediated gene flow in California cotton depends upon pollinator activity. Crop Sci. 45:1565–1570.
  • Schwember, A.R., and Bradford, K.J. 2005. Drying rates following priming affect temperature sensitivity of germination and longevity of lettuce seeds. HortScience 40: 778-781.
  • Alvarado, V., and Bradford, K.J. 2005. Hydrothermal time analysis of seed dormancy in true (botanical) potato seeds. Seed Sci. Res. 15: 77-88.
  • Bradford, K.J., Van Deynze, A., Gutterson, N., Parrott, W., Strauss, S.H. 2005. Regulating transgenic crops sensibly: lessons from plant breeding, biotechnology and genomics. Nature Biotechnol. 23: 439-444.
  • Sánchez, M.P., Gurusinghe, S.H., Bradford, K.J., and Vázquez-Ramos, J.M. 2005. Differential response of PCNA and CDK-A proteins and associated kinase activities to benzyladenine and abscisic acid during maize seed germination. J. Exp. Bot. 56: 515-523.
  • Bradford, K.J. 2005. Threshold models applied to seed germination ecology. New Phytol. 165: 338-341.


  • Bradford, K.J., and Alston, J.M. 2004. Horticultural biotechnology: challenges for commercial development. Chronica Horticulturae 44: 4-8.
  • Kende, H., Bradford, K.J., Brummell, D.A., Cho, H.T., Cosgrove, D.J., Fleming, A.J., Gehring, C., Lee, Y., McQueen-Mason, S., Rose, J.K.C., Voesenek, L.A.C.J. 2004. Nomenclature for members of the expansin superfamily of genes and proteins. Plant Mol. Biol. 55: 311-314.
  • Bradford, K.J., Alston, J.M, Lemaux, P.G. and Sumner, D.A. 2004. Challenges and opportunities for horticultural biotechnology. California Agriculture 58: 68-71. (Co-editor of entire issue)
  • Bradford, K.J., and Still, D.W. 2004. Applications of hydrotime analysis in seed testing. Seed Technology 26: 75-85.


  • Wu, C.T., and Bradford, K.J. 2003. Class I chitinase and b-1,3-glucanase are differentially regulated by wounding, methyl jasmonate, ethylene and gibberellin in tomato seeds and leaves. Plant Physiol. 133: 263-273.
  • Graff, G.D., Cullen, S.E., Bradford, K.J., Zilberman, D., Bennett, A.B. 2003. The public-private structure of intellectual property ownership in agricultural biotechnology. Nature Biotechnol. 21: 989-995.
  • Bradford, K.J., Downie, A.B., Gee, O.H., Alvarado, V.Y., Yang, H., Dahal, P. 2003. Abscisic acid and gibberellin differentially regulate expression of genes of the SNF1-related kinase complex in tomato seeds. Plant Physiol. 132: 1560-1576.
  • Whitmer, X., Nonogaki, H., Beers, E.P., Bradford, K.J., Welbaum, G.E. 2003. Characterization of chitinase activity and gene expression in muskmelon seeds. Seed Sci. Res. 13: 167-178.
  • Downie, B., Gurusinghe, S., Dahal, P., Thacker, R.R., Snyder, J.C., Nonogaki, H., Yim, K., Fukanaga, K., Alvarado, V., Bradford, K.J. 2003. Expression of a galactinol synthase gene in tomato seeds is up-regulated before maturation desiccation and again after imbibition whenever radicle protrusion is prevented. Plant Physiol. 131: 1347-1359.


  • Alvarado, V., and Bradford, K.J. 2002. A hydrothermal time model explains the cardinal temperatures for seed germination. Plant Cell Environ. 25: 1061-1069.
  • Gurusinghe, S., Powell, A.L.T., and Bradford, K.J. 2002. Enhanced expression of BiP is associated with treatments that extend storage longevity of primed tomato seeds. J. Amer. Soc. Hortic. Sci. 127: 528-534.
  • Bradford, K.J. 2002. Applications of hydrothermal time to quantifying and modeling seed germination and dormancy. Weed Sci. 50: 248–260.
  • Chen, F., Nonogaki, H., and Bradford, K.J. 2002. A gibberellin-regulated xyloglucan endotransglycosylase gene is expressed in the endosperm cap during tomato seed germination. J. Exp. Bot. 53: 215-223.


  • Kwong, F., Stodolski, L., Mari, J., Gurusinghe, S.H. and Bradford, K.J. 2001. Viability constants for delphinium and salvia seeds. Seed Technol. 23: 113-125.
  • Chen, F., Dahal, P., and Bradford, K.J. 2001. Two tomato expansin genes show divergent expression and localization in embryos during seed development and germination. Plant Physiol. 127 : 928-936.
  • Edelstein, M., Bradford, K.J. and Burger, D.W. 2001. Metabolic heat and CO2 production rates during germination of melon (Cucumis melo L.) seeds measured by microcalorimetry. Seed Sci. Res. 11: 265-272.
  • Wu, C.T., Leubner-Metzger, G., Meins, F. Jr. and Bradford K.J. 2001. Class I b-1,3-glucanase and chitinase are expressed specifically in the micropylar endosperm of tomato seeds prior to radicle emergence. Plant Physiol. 126: 1299-1313.
  • Gurusinghe, S., and Bradford, K.J. 2001. Galactosyl-sucrose oligosaccharides and potential longevity of primed seeds. Seed Sci. Res. 11: 121-133.


  • Chen, F., and Bradford, K.J. 2000. Expression of an expansin is associated with endosperm weakening during tomato seed germination. Plant Physiol. 124: 1265-1274.
  • Nonogaki, H., Gee, O.H., and Bradford, K.J. 2000. A germination-specific endo-b-mannanase gene is expressed in the micropylar endosperm cap of tomato seeds. Plant Physiol. 123: 1235-1245.


  • Cooley, M.B., Yang, H., Dahal, P. Mella, R.A., Downie, B., Haigh, A.M., and Bradford, K.J. 1999. Vacuolar H+-ATPase is expressed in response to gibberellin during tomato seed germination. Plant Physiol. 121: 1339-1347.
  • Sitrit, Y., Hadfield, K.A., Bennett, A.B., Bradford, K.J., and Downie, B. 1999. Expression of a polygalacturonase associated with tomato seed germination. Plant Physiol 121: 419-428.
  • Downie, B., Gurusinghe, S.H., and Bradford, K.J. 1999. Internal anatomy of individual tomato seeds: relationship to abscisic acid and germination physiology. Seed Sci. Res. 9: 117-128.
  • Cheng, Z., and Bradford, K.J. 1999. Hydrothermal time analysis of tomato seed germination responses to priming treatments. Exp. Bot. 50: 89-99.
  • Gurusinghe, S.H., Cheng, Z., and Bradford, K.J. 1999. Cell cycle activity during seed priming is not essential for germination advancement in tomato. J. Exp. Bot. 50: 101-106.


  • Downie, B., Dirk, L.M.A., Hadfield, K.A., Wilkins, T.A., Bennett, A.B., and Bradford, K.J. 1998. A gel diffusion assay for quantification of pectin methylesterase activity. Anal. Biochem. 264: 149-157.
  • Yim, K.-Y., and Bradford, K.J. 1998. Callose deposition is responsible for apoplastic semipermeability of the endosperm envelope of muskmelon seeds. Plant Physiol. 118: 83-90.
  • Still, D.W., and Bradford, K.J. 1998. Using hydrotime and ABA-time models to quantify seed quality of brassicas during development. J. Amer. Soc. Hort. Sci. 123: 692-699.
  • Bradford, K.J., and Cohn. MA. 1998. Seed biology and technology: At the crossroads and beyond. Seed Sci. Res. 8: 153-160.
  • Taylor, A.G., Allen, P.S., Bennett, M.A., Bradford, K.J., Burris, J.S., and Misra, M.K. 1998. Seed enhancements. Seed Sci. Res. 8: 245-256.
  • Welbaum, G.E., Bradford, K.J., Yim, K.-O., Booth, D.T., and Oluoch M.O. 1998. Biophysical, physiological and biochemical processes regulating seed germination. Seed Sci. Res. 8: 161-172.


  • Downie, B., Gurusinghe, S., Plopper, C., Bradford, K.J., Greenwood, J.S., and Bewley, J.D. 1997. Elongated cells adhering to the megagametophyte and sheathing the radicle of white spruce following completion of germination are derived from the embryo root cap. Int. J. Plant Sci.: 158: 738-746.
  • Dahal, P., Nevins, D.J., and Bradford, K.J. 1997. Relationship of endo-b-D-mannanase activity and cell wall hydrolysis in tomato endosperm to germination rates. Plant Physiol. 113: 1243-1252.
  • Still, D.W., Dahal, P., and Bradford, K.J. (1997) A single-seed assay for endo-b-mannanase activity from tomato endosperm and radicle tissues. Plant Physiol. 113: 13-20.
  • Still, D.W., and Bradford, K.J. (1997) Endo-b-mannanase activity from individual tomato endosperm caps and radicle tips in relation to germination rates. Plant Physiol. 113: 21-29.
  • Dutta, S., Bradford, K.J., and Nevins, D.J. 1997. Endo-b-mannanase activity present in cell wall extracts of lettuce (Lactuca sativa L.) endosperm prior to radicle emergence. Plant Physiol. 113: 155-161.


  • Dahal, P., N.-S. Kim, and K.J. Bradford. 1996. Respiration and germination rates of tomato seeds at suboptimal temperatures and reduced water potentials. J. Exp. Bot. 47: 941-947.


  • Baker, E.H., K.J. Bradford, J.A. Bryant, and T.L. Rost. 1995. A comparison of desiccation-related proteins (dehydrin and QP47) in peas (Pisum sativum). Seed Sci. Res. 5: 185-193.


  • Bradford, K.J., and A.J. Trewavas. 1994. Sensitivity thresholds and variable time scales in plant hormone action. Plant  Physiol 105: 1029-1036.
  • Baker, E.H., and K.J. Bradford. 1994. The fluorescence assay for Maillard product accumulation does not correlate with seed viability. Seed Sci. Res. 4: 103-108.
  • Berjak, P., K.J. Bradford, D.A. Kovach, and N.W. Pammenter. 1994. Differential effects of temperature on ultrastructural responses to dehydration in seeds of Zizania palustris. Seed Sci. Res. 4: 111-122.
  • Bradford, K.J., and A.M. Haigh. 1994. Relationship between accumulated hydrothermal time during seed priming and subsequent seed germination rates. Seed Sci. Res. 4: 63-70.
  • Dahal, P., and K.J. Bradford. 1994. Hydrothermal time analysis of tomato seed germination at suboptimal temperature and reduced water potential. Seed Sci. Res. 4: 71-80.
  • Bradford, K.J., and O.A. Somasco. 1994. Water relations of lettuce seed thermoinhibition. I. Priming and endosperm effects on base water potential. Seed Sci. Res. 4: 1-10.
  • Dutta, S., and K.J. Bradford. 1994. Water relations of lettuce seed thermoinhibition. II. Ethylene and endosperm effects on base water potential. Seed Sci. Res. 4: 11-18.
  • Dutta, S., K.J. Bradford, and D.J. Nevins. 1994. Cell-wall autohydrolysis in isolated endosperms of lettuce (Lactuca sativa L.). Plant Physiol. 104: 623-628.
  • Still, D.W, D.A. Kovach, and K.J. Bradford. 1994. Development of desiccation tolerance during embryogenesis in rice (Oryza sativa) and wild rice (Zizania palustris): dehydrin expression, abscisic acid content, and sucrose accumulation. Plant Physiol. 104: 431-438.
  • Bradford, K.J. 1994. Water stress and the water relations of seed development: a critical review. Crop Sci. 34: 1-11.


  • Bradford, K.J., A.M. Tarquis, and J.M. Durán. 1993. A population-based threshold model describing the relationship between germination rates and seed deterioration. J. Exp. Bot. 264: 1225-1234.
  • Ni, B.-R., and K.J. Bradford. 1993. Germination and dormancy of abscisic acid- and gibberellin-deficient mutant tomato  seeds. Sensitivity of germination to abscisic acid, gibberellin, and water potential. Plant Physiol. 101: 607-617.


  • Tarquis, A. M., and K. J. Bradford. 1992. Prehydration and priming treatments that advance germination also increase the  rate of deterioration of lettuce seeds. J. Exp. Bot. 43: 307-317.
  • Ni, B.R, and K.J. Bradford. 1992. Quantitative models characterizing seed germination responses to abscisic acid and osmoticum. Plant Physiol. 98: 1057-1068.
  • Kovach, D.A., and K.J. Bradford 1992. Temperature dependence of viability and dormancy of Zizania palustris var. interior seeds stored at high moisture contents. Ann. Bot. 69: 297-301.
  • Kovach, D.A., and K.J. Bradford. 1992. Imbibitional damage and desiccation tolerance of wild rice (Zizania palustris) seeds. Exp. Bot. 43: 747-757.
  • Bradford, K.J., and P.M. Chandler. 1992. Expression of ‘dehydrin-like’ proteins in embryos and seedlings of Zizania palustris and Oryza sativa during dehydration. Plant Physiol. 99: 488-494.


  • Welbaum, G.E., and K.J. Bradford. 1991. Water relations of seed development and germination in muskmelon (Cucumis melo L.). VII. Influence of afterripening and ageing on germination responses to temperature and water potential. J. Exp. Bot. 42: 1137-1145.
  • Welbaum, G.E., and K.J. Bradford. 1991. Water relations of seed development and germination in muskmelon (Cucumis melo L.). VI. Influence of priming on germination responses to temperature and water potential during seed development. J. Exp. Bot. 42: 393-399.
  • Mayberry, K.S., K.J. Bradford, and V.E. Rubatzky. 1991. Yellow cotyledon: a seedling disorder of broccoli. HortScience 26: 21-23.


  • Bradford, K.J. 1990. A water relations analysis of seed germination rates. Plant Physiol. 94: 840-849.
  • Welbaum, G.E., and K.J. Bradford. 1990. Water relations of seed development and germination in muskmelon (Cucumis melo L.). III. Sensitivity of germination to water potential and abscisic acid during development. Plant Physiol. 92: 1029-1037. Welbaum, G.E., and K.J. Bradford. 1990. Water relations of seed development and germination in muskmelon (Cucumis melo L.). IV. Characteristics of the perisperm during seed development. Plant Physiol. 92: 1038-1045.
  • Welbaum, G.E., and K.J. Bradford. 1990. Water relations of seed development and germination in muskmelon (Cucumis melo L.). V. Water relations of imbibition and germination. Plant Physiol. 92: 1046-1052.
  • Dahal, P., K.J. Bradford, and R.A. Jones. 1990. Effects of priming and endosperm integrity on seed germination rates of tomato genotypes. I. Germination at suboptimal temperature. J. Exp. Bot. 41: 1431-1439.
  • Dahal, P., and K.J. Bradford. 1990. Effects of priming and endosperm integrity on seed germination rates of tomato genotypes. II. Germination at reduced water potential. J. Exp. Bot. 41: 1441-1453.
  • Trawatha,  S.E.,  J.J.  Steiner,  and  K.J.  Bradford. 1990. Laboratory vigor tests used to predict pepper seedling field emergence performance. Crop Sci. 30: 713-717.
  • Bradford, K.J., J.J. Steiner, and S.E. Trawatha. 1990. Seed priming influence on germination and emergence of pepper seed lots. Crop Sci. 30: 718-721.
  • Argerich, C.A., K.J. Bradford, and F.M. Ashton. 1990. Influence of seed vigor and preplant herbicides on the emergence, seedling growth and yield of tomato. HortScience 25: 288-291.
  • Kelly, M.O., and K.J. Bradford. 1990. Ethylene synthesis and growth of tomato hypocotyls: induction by auxin and fusicoccin and inhibition by vanadate. J. Plant Growth Regul. 9: 43-49.


  • Welbaum, G.E., and K.J. Bradford. 1989. Water relations of seed development and germination in muskmelon (Cucumis melo L.). II. Development of germinability, vigour, and desiccation tolerance. J. Exp. Bot. 40: 1355-1362.
  • Ursin, V.M., and K.J. Bradford. 1989. Auxin and ethylene regulation of petiole epinasty in two developmental mutants of tomato, diageotropica and Epinastic. Plant Physiol. 90: 1341-1346.
  • Ursin, V.M., and K.J. Bradford. 1989. A unique phenotype in heterozygotes of the auxin-insensitive mutant of tomato, diageotropica. Plant Physiol. 90: 1243-1245.
  • Argerich, C.A., K.J. Bradford and A.M. Tarquis. 1989. The effects of priming and ageing on resistance to deterioration of tomato seeds. J. Exp. Bot. 40: 593-598.
  • Argerich, C.A., and K.J. Bradford. 1989. The effects of priming and ageing on seed vigour in tomato. J. Exp. Bot. 40: 599-607.
  • Fujino, D.W., D.W. Burger and K.J. Bradford. 1989. Ineffectiveness of ethylene biosynthetic and action inhibitors to phenotypically revert the Epinastic mutant of tomato (Lycopersicon esculentum Mill.). J. Plant Growth Regul. 8:53-61.


  • Fujino, D.W., S.J. Nissen, A.D. Jones, D.W. Burger and K.J. Bradford. 1988. Quantification of indole-3-acetic acid in dark- grown seedlings of the diageotropica and Epinastic mutants of tomato (Lycopersicon esculentum Mill.). Plant Physiol. 88:780-784.
  • Fujino, D.W., D.W. Burger, S.F. Yang and K.J. Bradford. 1988. Characterization of an ethylene overproducing mutant of tomato (Lycopersicon esculentum Mill. cv. VFN8). Plant Physiol. 88:774-779.
  • Alvarado, A.D., and K.J. Bradford. 1988. Priming and storage of tomato (Lycopersicon lycopersicum) seeds.  II.  Influence of  a second priming treatment after storage on germination and field emergence. Seed. Sci. & Technol. 16:613-623.
  • Alvarado, A.D., and K.J. Bradford. 1988. Priming and storage of tomato (Lycopersicon lycopersicum) seeds. I. Effects of storage temperature on germination rate and viability. Seed Sci. & Technol. 16:601-612.
  • Bradford, K.J., D.M. May, B.J. Hoyle, Z.S. Skibinski, S.J. Scott and K.B. Tyler. 1988. Seed and soil treatments to improve emergence of muskmelon from cold or crusted soils. Crop Sci. 28:1001-1005.
  • Noland, T.L., J.D. Maguire, R.N. Oliva, K.J. Bradford, J.L. Nelson, D. Grabe, and S. Currans. 1988. Effect of plant density on carrot seed yield and quality under seed-to-seed production systems in California, Oregon, and Washington. J. Applied Seed Prod. 6: 36-43.
  • Oliva, R.N., T. Tissaoui and K.J. Bradford. 1988. Relationships of plant density and harvest index to seed yield and quality in carrot. J. Amer. Soc. Hort. Sci. 113:532-537.
  • Welbaum, G.E., and K.J. Bradford. 1988. Water relations of seed development and germination in muskmelon (Cucumis melo). I. Water relations of seed and fruit development. Plant Physiol. 86:406-411.


  • Kwon, O.-S., and K.J. Bradford. 1987. Tomato seed development and quality as influenced by preharvest treatment with ethephon. HortScience 22:588-591
  • Alvarado,  A.D.,  K.J.  Bradford  and  J.D.  Hewitt. 1987. Osmotic  priming  of tomato seeds: Effects on germination, field emergence, seedling growth and fruit yield. J. Amer. Soc. Hort. Sci. 112:427-432.
  • Valdes, V.M., and K.J. Bradford. 1987. Effects of seed coating and osmotic priming on the germination of lettuce seeds at  high temperatures. J. Amer. Soc. Hort. Sci. 112:153-156.


  • Kelly, M.O., and K.J. Bradford. 1986. Insensitivity of the diageotropica tomato mutant to auxin. Plant Physiol. 82:713-717. Bradford, K.J. 1986. Manipulation of seed water relations via osmotic priming to improve germination under stress conditions. HortScience 21:1105-1112.
  • Eisinger, B.A., and K.J. Bradford. 1986. Role of seed mineral content in the occurrence of transverse cotyledon cracking of snap bean (Phaseolus vulgaris L.). J. Amer. Soc. Hort. Sci. 111:110-113.


  • Valdes, V.M., K.J. Bradford and K.S. Mayberry. 1985. Alleviation of thermodormancy in coated lettuce seeds by seed priming. HortScience 20(6):1112-1114.


  • Kwon, O.-S., and K.J. Bradford. 1984. Seed quality and fruit ripening of tomato (Lycopersicon esculentum Mill.) as influenced by preharvest treatment with ethephon. J. Appl. Seed Production 2: 54-57.


  • Bradford, K.J. 1983. Involvement of plant growth substances in the alteration of leaf gas exchange of flooded tomato plants. Plant Physiol. 73:480-483.
  • Bradford, K.J. 1983. Effects of soil flooding on leaf gas exchange of tomato plants. Plant Physiol. 73:475-479.
  • Bradford, K.J. 1983. Water relations and growth of the flacca tomato mutant in relation to abscisic acid. Plant Physiol. 72: 251-255.
  • Bradford, K.J., T.D. Sharkey and G.D. Farquhar. 1983. Gas exchange, stomatal behavior and d13C values of the flacca tomato mutant in relation to abscisic acid. Plant Physiol. 71:245-250.


  • Bradford, K.J., T.C. Hsiao and S.F. Yang. 1982. Inhibition of ethylene synthesis in tomato plants subjected to anaerobic root stress. Plant Physiol 70:1503-1507.
  • Bradford, K.J., and T.C. Hsiao. 1982. Stomatal behavior and water relations of waterlogged tomato plants. Plant Physiol. 70:1508-1513.


  • Bradford, K.J., and S.F. Yang. 1981. Physiological responses of plants to waterlogging. HortScience 16:25-30.


  • Bradford,  K.J.,  and  S.F.  Yang. 1980. Stress-induced ethylene production in the ethylene-requiring tomato mutant diageotropica. Plant Physiol. 65:327-330.
  • Bradford, K.J., and S.F. Yang. 1980. Xylem transport of 1-amino-cyclopropane-1-carboxylic acid, an ethylene precursor, in waterlogged tomato plants. Plant Physiol. 65:322-326.


  • Saltveit, M.E., Jr., K.J. Bradford, and D.R. Dilley. 1978. Silver ion inhibits ethylene synthesis and action in ripening fruits. J. Amer. Soc. Hort. Sci. 103(4):427-475.
  • Bradford, K.J., and D.R. Dilley. 1978. Effects of root anaerobiosis on ethylene production, epinasty, and growth of tomato plants. Plant Physiol. 61:506-509.